In The Crew for the PS4; The story begins with the main character Alex Taylor being pursued by local law enforcement near Detroit. During the chase, he receives a call from Harry who wants him to participate in a race in the city. After losing the cops, he finds the Camaro that Harry loans him. Harry explains to him that Alexís older brother Dayton, the founder of the 5-10 motor club, wants to speak with him. Alex participates and wins the race. After his win, Dayton arrives and orders Alex to drive him to the bridge. Once there, Dayton tells him to keep his head down. A vehicle pulls up, Dayton goes and talks with the driver before walking back to the Camaro; however before he can get there, the driver shoots him and drives off.
The police restrain Alex as Dayton dies. Alex is charged and later convicted of Daytonís murder by FBI Special Agent Coburn and is sent to prison. 5 years later, an FBI agent named Zoe finds Alex and informs him that the new leader of the 5-10 motor club, a man named Shiv was the one who killed Dayton and that Coburn in collaboration with Shiv framed Alex for Dayton's murder. Zoe offers Alex a chance to get Shiv behind bars if he helps her get evidence against Special Agent Coburn, Alex takes the offer. After getting released from prison and beating Zoe in a race, Alex accepts his first mission to help a 5-10 named Troy. After doing a series of missions for him, Alex is sent to St. Louis to kill a 5-10 of higher rank, called a V2.
At a stage in the game Alex goes to New York and is reunited with Harry, who agrees to help him with his mission. Alex gets a dirt car and meets with Eric, racing him and later doing missions for him. Meanwhile, Zoe and Alex are suspicious of Harry when he gets secretive. It is later revealed that Harry was having affairs with Dayton's girlfriend, Connie, and stealing cars that Alex and Eric were forced to recover. Alex also has to deliver something to a mysterious person, later revealed to be Coburn's. The last mission in New York is three parts: Alex races against someone with a better car, who says that the winner will get the loser's car. Furious after losing, the driver calls the cops, but Alex manages to escape.